Holotheria - Biblioteka.sk

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Holotheria
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Over 70% of mammal species are in the orders Rodentia (blue), Chiroptera (red) and Soricomorpha (yellow).
  Pilosa

Mammalia is a class of animal within the phylum Chordata. Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is universally accepted; McKenna & Bell (1997) and Wilson & Reader (2005) provide useful recent compendiums.[1] Many earlier ideas from Linnaeus et al. have been completely abandoned by modern taxonomists, among these are the idea that bats are related to birds or that humans represent a group outside of other living things.[2] Competing ideas about the relationships of mammal orders do persist and are currently in development. Most significantly in recent years, cladistic thinking has led to an effort to ensure that all taxonomic designations represent monophyletic groups. The field has also seen a recent surge in interest and modification due to the results of molecular phylogenetics.

George Gaylord Simpson's classic "Principles of Classification and a Classification of Mammals" (Simpson, 1945) taxonomy text laid out a systematics of mammal origins and relationships that was universally taught until the end of the 20th century.

Since Simpson's 1945 classification, the paleontological record has been recalibrated, and the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself, partly through the new concept of cladistics. Though field work gradually made Simpson's classification outdated, it remained the closest thing to an official classification of mammals. See List of placental mammals and List of monotremes and marsupials for more detailed information on mammal genera and species.

Molecular classification of placentals

Molecular studies by molecular systematists, based on DNA analysis, in the early 21st century have revealed new relationships among mammal families. Classification systems based on molecular studies reveal three major groups or lineages of placental mammals, Afrotheria, Xenarthra, and Boreotheria. which diverged from early common ancestors in the Cretaceous.[3]

The relationships between these three lineages are contentious, and all three have been proposed as basal in different hypotheses.[3][4][5]

The following taxonomy only includes living placentals (infraclass Eutheria):[citation needed]

Atlantogenata

Afrotheria

Xenarthra

Boreoeutheria

Euarchontoglires

Masoala fork-marked lemur (Cheirogaleus) Phaner furcifer
  • Superorder Euarchonta
  • Superorder Glires
    • Order Lagomorpha: pikas, rabbits, hares (Eurasia, Africa, Americas)
      Arctic hare (Leporid)
      • Family Leporidae: (60 species), rabbits and hares (Eurasia, Africa, Americas)
      • Family Ochotonidae: (30 species), pikas (Holarctic)
    • Order Rodentia: rodents (cosmopolitan)
      • Suborder Castorimorpha
        • Family Castoridae: (2 species) beavers (Holarctic)
        • Family Geomyidae: (about 35 species) pocket gophers (North America)
        • Family Heteromyidae: (about 59 species) kangaroo rats and kangaroo mice (North America)
      • Suborder Myomorpha
        • Family Dipodidae: (33 species) jerboas (Africa, Eurasia, North America)
        • Family Zapodidae: (11 species) jumping mice (North America, Asia)
        • Family Sicistidae: (19 species) birch mice (Eurasia)
        • Family Platacanthomyidae: (3 species) spiny dormouse (Southeast Asia)
        • Family Spalacidae: (37 species) zokors, root rats, blind mole rats (Africa, Eurasia)
        • Family Calomyscidae: (8 species) mouse-like hamsters (Asia)
        • Family Nesomyidae: (68 species) old endemic African muroids (Africa, Madagascar)
        • Family Cricetidae: (about 580 species) hamsters, voles, and New World rats and mice (Holarctic, South America)
        • Family Muridae: (about 1,383 species) Old World rats and mice and gerbils (Africa, Eurasia, Australia)
      • Suborder Anomaluromorpha
        • Family Anomaluridae: (6 species) scaly-tailed flying squirrels (Africa)
        • Family Pedetidae: (2 species) springhares or springhaas (Africa)
      • Suborder Hystricomorpha
        • Family Ctenodactylidae: (5 species) gundis (Africa, Asia)
        • Family Diatomyidae: (1 species) Laotian rock rat (Southeast Asia)
        • Family Hystricidae: (11 Species) Old World porcupines (Africa, Asia)
        • Family Bathyergidae: (about 21 species) African mole-rats (Africa)
        • Family Petromuridae: (1 species) rock dassies (Africa)
        • Family Thryonomyidae: (2 species) cane rats (Africa)
        • Family Erethizontidae: (19 species) New World porcupines (New World)
        • Family Chinchillidae: (3 species) chinchillas and viscachas (South America)
        • Family Dinomyidae: (1 species) pacarana (South America)
        • Family Caviidae: (18 species) cavies and capybara (South America)
        • Family Dasyproctidae: (13 species) agoutis and acouchis (South America)
        • Family Cuniculidae: (about 3 species) paca (South America)
        • Family Ctenomyidae: (about 60 species) tuco-tucos (South America)
        • Family Octodontidae: (14 species) degus (South America)
        • Family Abrocomidae: (9 species) chinchilla-rats (South America)
        • Family Echimyidae: spiny rats (South America)
        • Family Capromyidae: (10 species) hutias (South America)
        • Family Heptaxodontidae: giant Hutias (recently extinct)
        • Family Myocastoridae: (57 species) nutrias (South America)
      • Suborder Sciuromorpha
        Mountain beaver (Aplodont)
        • Family Aplodontiidae: (1 species) mountain beaver (western North America)
        • Family Sciuridae: (about 285 species) squirrels, chipmunks, and marmots (cosmopolitan except Australia)
        • Family Gliridae: (29 species) dormice (Africa, Eurasia)

Laurasiatheria

  • Order Eulipotyphla
    • Family Solenodontidae: (2 species) solenodons (Cuba, Hispaniola)
    • Family Nesophontidae: nesophontes (West Indies shrews) (recently extinct)
    • Family Soricidae: (385 species) shrews (Eurasia, Africa, North America to northern South America)
    • Family Talpidae: (59 species) moles, shrew-moles, desmans (Eurasia, North America)
    • Family Erinaceidae: (26 species) hedgehogs, gymnures (Eurasia, Africa)
    • Family Galericidae: (8 species) moonrats (southeast Asia)
  • Grandorder Chiroptera
    • Order Chiroptera: bats
      • Suborder Yinpterochiroptera
        • Family Pteropodidae: (about 197 species) flying foxes (Africa, Eurasia, Australia)
        • Family Hipposideridae: (84 species) trident bats, leaf-nosed bats
        • Family Rhinolophidae: (106 species) horseshoe bats (Old World)
        • Family Rhinopomatidae: (6 species) mouse-tailed bats (Africa, Southeast Asia)
        • Family Craseonycteridae: (1 species) Kitti's hog-nosed bat (Thailand)
        • Family Megadermatidae: (6 species) false vampire bats (Africa, Southeast Asia, Australia)
      • Suborder Yangochiroptera
        • Family Emballonuridae: (54 species) sac-winged bats (southern continents)
        • Family Nycteridae: (about 15 species) slit-faced bats (Africa, Southeast Asia)
        • Family Mystacinidae: (about 2 species) short-tailed bats (New Zealand)
        • Family Thyropteridae: (5 species) sucker-footed bats (South America)
        • Family Furipteridae: (2 species) smoky bats (South America)
        • Family Noctilionidae: (2 species) fishing bats (South America)
        • Family Mormoopidae: (about 11 species) leaf-chinned bats (South America)
        • Family Phyllostomidae: (192 species) leaf-nosed bats (South America)
        • Family Myzopodidae: (2 species) sucker-footed bats (Madagascar)
        • Family Natalidae: (10 species) funnel-eared bats (South America)
        • Family Molossidae: (about 110 species) free-tailed bats (cosmopolitan)
        • Family Miniopteridae: (about 40 species) long-fingered bats (Africa, Eurasia, Australia)
        • Family Cistugidae: (2 species) wing-gland bats (Southern Africa)
        • Family Vespertilionidae: (over 300 species) vesper bats (cosmopolitan)
  • Grandorder Ferae
    • Order Pholidota
      • Family Manidae: (about 8 species) pangolins, scaly anteaters (Africa, South Asia)
    • Order Carnivora: carnivorans (cosmopolitan)
      • Suborder Feliformia
        • Family Nandiniidae: (4 species) African palm civet (Central Africa)
        • Family Prionodontidae: (2 species) Asiatic linsangs (Southeast Asia)
        • Family Felidae: (41 species) cats (cosmopolitan except Australia)
        • Family Viverridae: (33 species) civets, Asiatic palm civets (Africa, Southern Europe, Southeast Asia)
        • Family Herpestidae: (34 species) mongooses (Africa, Asia, Southern Europe)
        • Family Eupleridae: (10 species) Malagasy carnivorans (Madagascar)
        • Family Hyaenidae: (4 species) hyaenas, aardwolf (Africa)
      • Suborder Caniformia
        • Family Canidae: (38 species) dogs (cosmopolitan)
        • Family Ursidae: (8 species) bears (Europe, Asia, New World)
        • Family Otariidae: (15 species) eared seals (cosmopolitan except North Atlantic)
        • Family Odobenidae: (1 species) walrus (Northern North American, Northern Europe, Northern Asia)
        • Family Phocidae: (18 species) true seals (cosmopolitan)
        • Family Ailuridae: (1 species) red panda (South-Central Asia)
        • Family Mephitidae: (12 species) skunks (Southeast Asia, New World)
        • Family Mustelidae: (about 69 species) weasels and relatives (cosmopolitan except Australia)
        • Family Procyonidae: (14 species) ringtails, olingos, kinkajou, raccoons, coatis (New World)
  • Grandorder Euungulata
    • Order Perissodactyla: odd-toed ungulates
      • Family Equidae: (13 species) horses, zebras, donkeys (Africa, West and Central Asia)
      • Family Tapiridae: (3 species) tapirs (Central and South America, Southeast Asia)
      • Family Rhinocerotidae: (5 species) rhinoceroses (Africa, Southeast Asia)
    • Order Artiodactyla: even-toed ungulates (now includes Cetaceans)
      • Suborder Suiformes
        • Family Suidae: (18 species) pigs (Africa, Eurasia)
          Pig and piglet
        • Family Tayassuidae: (about 3 species) peccaries (New World)
      • Suborder Tylopoda
        • Family Camelidae: (7 species) camels (South America, Asia)
      • Suborder Ruminantia
        • Family Tragulidae: (10 species) mouse-deer (Africa, Asia)
        • Family Antilocapridae: (1 species) pronghorn (North America)
        • Family Giraffidae: (2-9 species) giraffe and okapi (Africa)
          Giraffe
        • Family Cervidae: (26 species) deer (Holarctic, South America)
        • Family Moschidae: (7 species) musk deer (Asia)
          Muntjac deer
        • Family Bovidae: (143 species) cattle, antelope, sheep, etc. (Africa, Holarctic)
          Pair of Icelandic Sheep
      • Suborder Whippomorpha
        • Family Hippopotamidae: (2 species) hippos (Africa)
        • Infraorder Cetacea
          • Parvorder Mysticeti
            • Family Balaenopteridae: (10 species) rorquals and grey whales (cosmopolitan)
            • Family Balaenidae: (4 species) right and bowhead whales (polar and temperate waters)
            • Family Eschrichtiidae: (1 species) gray whale (North Pacific and North Atlantic)
            • Family Neobalaenidae: (1 species) pygmy right whales (southern hemisphere)
          • Parvorder Odontoceti
            • Family Delphinidae: (about 37 species) dolphins (cosmopolitan)
            • Family Monodontidae: (2 species) beluga and narwhal (Arctic, Atlantic, and Pacific)
            • Family Phocoenidae: (8 species) porpoises (cosmopolitan)
            • Family Physeteridae: (3 species) sperm whales (cosmopolitan)
            • Family Kogiidae: (2 species) dwarf sperm whales (cosmopolitan)
            • Family Platanistidae: (2 species) South Asian river dolphin (Southern Asia)
            • Family Iniidae: (1-4 species) Amazon River dolphin (South America)
            • Family Pontoporiidae: (1 species) La Plata River dolphin (South America)
            • Family Lipotidae: baiji
            • Family Ziphiidae: (24 species) beaked whales (cosmopolitan)

Standardized textbook classification

A somewhat standardized classification system has been adopted by most current mammalogy classroom textbooks. The following taxonomy of extant and recently extinct mammals is taken from the 6th edition of Vaughan's Mammalogy.[1] This approach emphasizes an initial split between egg-laying prototherians and live-bearing therians. The therians are further divided into the marsupial Metatheria and the "placental" Eutheria. No attempt is made in this classification to further distinguish among the orders within these subclasses and infraclasses. This system also makes no note of the position of entirely fossil groups.

In this and later taxonomies, families are merely listed under the order to which they belong. More detailed relationships among families is presented in the article of each order.

Subclass Prototheria

Subclass Theria

McKenna/Bell classification

In 1997, the classification of mammals was revised by Malcolm C. McKenna and Susan K. Bell.[10] The Classification of Mammals Above the species level, here referred to as the "McKenna/Bell classification", is a comprehensive work on the systematics, relationships, and occurrences of all mammal taxa, living and extinct, down through the rank of genus. The authors worked together as paleontologists at the American Museum of Natural History, New York. McKenna inherited the project from Simpson and, with Bell, constructed a completely updated hierarchical system, covering living and extinct taxa that reflects the historical genealogy of Mammalia.

The McKenna/Bell hierarchical listing of all of the terms used for mammal groups above the species includes extinct mammals as well as modern groups, and introduces some fine distinctions such as legions and sublegions and ranks which fall between classes and orders that are likely to be glossed over by the layman.

Click on the highlighted link for a table comparing the traditional and the new McKenna/Bell classifications of mammals.

Extinct groups are represented by †.

Subclass Prototheria

(monotremes)

Subclass Theriiformes







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